van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
アジメドジョウ
本村, 2020 (JAF list ver. 1 - 6); 本村, 2021 (JAF list ver. 7 - 12); 本村, 2022 (JAF list ver. 13 - 18); 本村, 2023 (JAF list ver. 19 - 21)
Niwaella delicata
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.; Kottelat, 2012Kottelat, M. 2012. CONSPECTUS COBITIDUM: AN INVENTORY OF THE LOACHES OF THE WORLD (TELEOSTEI: CYPRINIFORMES: COBITOIDEI). THE RAFFLES BULLETIN OF ZOOLOGY, Supplement 26: 1–199.; van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; Perdices et al., 2016Perdices, A., Bohlen, J., Šlechtová, V. & Doadrio, I. 2016. Molecular Evidence for Multiple Origins of the European Spined Loaches (Teleostei, Cobitidae). PLoS ONE, 11(1): e0144628. DOI: 10.1371/journal.pone.0144628 URL; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; Chen et al., 2017Chen, Y., He, D., Chen, H. & Chen, Y. 2017. Taxonomic study of the genus Niwaella (Cypriniformes: Cobitidae) from East China, with description of four new species. Zoological Systematics, 42(4): 490-507. DOI: 10.11865/zs.201723 URL
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Nakajima et al., 2008Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. 2008. Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57: 10-15.
striata complex Kyushu form
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
striata complex Small race, Kyushu form
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
magno = ラテン語で「大きい」の意味.本種は日本産スジシマドジョウ種群の中でもっとも大型になることにちなむ
学名・和名の変遷β
学名
和名
出典
Cobitis magnostriata
Yang et al., 2015Yang, L., Sado, T., Hirt, M.V., Pasco-Viel, E., Arunachalam, M., Li, J., Wang, X., Freyhof, J., Saitoh, K., Simons, A.M., Miya, M., He, S. & Mayden, R.L. 2015. Phylogeny and polyploidy: Resolving the classification of cyprinine fishes (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution, 85: 97-116.; van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; Tabata et al., 2016Tabata, R., Kakioka, R., Tominaga, K., Komiya, T. & Watanabe, K. 2016. Phylogeny and historical demography of endemic fishes in Lake Biwa: the ancient lake as a promoter of evolution and diversification of freshwater fishes in western Japan. Ecology and Evolution, DOI: 10.1002/ece3.2070 URL; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; Chen et al., 2017Chen, Y., He, D., Chen, H. & Chen, Y. 2017. Taxonomic study of the genus Niwaella (Cypriniformes: Cobitidae) from East China, with description of four new species. Zoological Systematics, 42(4): 490-507. DOI: 10.11865/zs.201723 URL; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL; Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
Kottelat, 2012Kottelat, M. 2012. CONSPECTUS COBITIDUM: AN INVENTORY OF THE LOACHES OF THE WORLD (TELEOSTEI: CYPRINIFORMES: COBITOIDEI). THE RAFFLES BULLETIN OF ZOOLOGY, Supplement 26: 1–199.; Chen et al., 2017Chen, Y., He, D., Chen, H. & Chen, Y. 2017. Taxonomic study of the genus Niwaella (Cypriniformes: Cobitidae) from East China, with description of four new species. Zoological Systematics, 42(4): 490-507. DOI: 10.11865/zs.201723 URL
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Cobitis sp. ‘yamato’ complex
Nakajima et al., 2011Nakajima, J., Ohata, T. & Onikura, N. 2011. Longitudinal distribution patterns of three spined loach species (Cobitidae, Cobitis) in the Onga River system, northern Kyushu Island, Japan. Folia Zoologica, 60: 322–327.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
yamato complex
Nakajima et al., 2008Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. 2008. Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57: 10-15.
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; Chen et al., 2017Chen, Y., He, D., Chen, H. & Chen, Y. 2017. Taxonomic study of the genus Niwaella (Cypriniformes: Cobitidae) from East China, with description of four new species. Zoological Systematics, 42(4): 490-507. DOI: 10.11865/zs.201723 URL; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
striata complex Small race, Sanyo form
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; Tabata et al., 2016Tabata, R., Kakioka, R., Tominaga, K., Komiya, T. & Watanabe, K. 2016. Phylogeny and historical demography of endemic fishes in Lake Biwa: the ancient lake as a promoter of evolution and diversification of freshwater fishes in western Japan. Ecology and Evolution, DOI: 10.1002/ece3.2070 URL; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; Chen et al., 2017Chen, Y., He, D., Chen, H. & Chen, Y. 2017. Taxonomic study of the genus Niwaella (Cypriniformes: Cobitidae) from East China, with description of four new species. Zoological Systematics, 42(4): 490-507. DOI: 10.11865/zs.201723 URL; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
striata complex Small race, Biwa form
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
striata complex Spotted small race
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; Chen et al., 2017Chen, Y., He, D., Chen, H. & Chen, Y. 2017. Taxonomic study of the genus Niwaella (Cypriniformes: Cobitidae) from East China, with description of four new species. Zoological Systematics, 42(4): 490-507. DOI: 10.11865/zs.201723 URL; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
striata complex Tokai small race
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
striata complex Yodo small race
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
オオヨドシマドジョウ
☆Nakajima & Suzawa, 2015Nakajima, J. & Suzawa, Y. 2015. Cobitis sakahoko, a new species of spined loach (Cypriniformes: Cobitidae) from southern Kyushu Island, Japan. Ichthyological Research, 63: 68–78. DOI: 10.1007/s10228-015-0476-5. URL; 細谷ほか, 2015細谷和海・内山りゅう・藤田朝彦・武内啓明・川瀬成吾. 2015. 山渓ハンディ図鑑15 日本の淡水魚. 山と渓谷社,528pp. Amazon; 中島・内山, 2017中島 淳・内山りゅう. 2017. 日本のドジョウ 形態・生態・文化と図鑑. 山と渓谷社,223pp. Amazon; 中坊編, 2018中坊徹次(編). 2018. 小学館の図鑑Z 日本魚類館. 小学館,xvi+524pp. Amazon; 細谷ほか, 2019細谷和海・内山りゅう・藤田朝彦・武内啓明・川瀬成吾. 2019. 山渓ハンディ図鑑15 増補改訂 日本の淡水魚. 山と渓谷社, 560pp. Amazon; 本村, 2020 (JAF list ver. 1 - 6); 本村, 2021 (JAF list ver. 7 - 12); 本村, 2022 (JAF list ver. 13 - 18); 本村, 2023 (JAF list ver. 19 - 21)
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.; Kottelat, 2012Kottelat, M. 2012. CONSPECTUS COBITIDUM: AN INVENTORY OF THE LOACHES OF THE WORLD (TELEOSTEI: CYPRINIFORMES: COBITOIDEI). THE RAFFLES BULLETIN OF ZOOLOGY, Supplement 26: 1–199.; Perdices et al., 2016Perdices, A., Bohlen, J., Šlechtová, V. & Doadrio, I. 2016. Molecular Evidence for Multiple Origins of the European Spined Loaches (Teleostei, Cobitidae). PLoS ONE, 11(1): e0144628. DOI: 10.1371/journal.pone.0144628 URL; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
ヒナイシドジョウ
☆★Suzawa, 2006Suzawa, Y. 2006. A new loach, Cobitis shikokuensis (Teleostei: Cobitidae), from Shikoku Island, Japan. Ichthyological Research, 53: 315-322. DOI: 10.1007/s10228-006-0352-4 URL; 中島ほか, 2012中島 淳・洲澤 譲・清水孝昭・斉藤憲治. 2012. 日本産シマドジョウ属魚類の標準和名の提唱. 魚類学雑誌, 59(1): 86-95.; 中坊編, 2013中坊徹次(編). 2013. 日本産魚類検索 全種の同定 第三版. 東海大学出版会,l+2430pp. URL; van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; 環境省, 2015環境省. 2015. レッドデータブック2014 日本の絶滅のおそれのある野生生物 4 汽水・淡水魚類. ぎょうせい,xxx+414pp. Amazon; 細谷ほか, 2015細谷和海・内山りゅう・藤田朝彦・武内啓明・川瀬成吾. 2015. 山渓ハンディ図鑑15 日本の淡水魚. 山と渓谷社,528pp. Amazon; 中島・内山, 2017中島 淳・内山りゅう. 2017. 日本のドジョウ 形態・生態・文化と図鑑. 山と渓谷社,223pp. Amazon; 中坊編, 2018中坊徹次(編). 2018. 小学館の図鑑Z 日本魚類館. 小学館,xvi+524pp. Amazon; 細谷ほか, 2019細谷和海・内山りゅう・藤田朝彦・武内啓明・川瀬成吾. 2019. 山渓ハンディ図鑑15 増補改訂 日本の淡水魚. 山と渓谷社, 560pp. Amazon; 本村, 2020 (JAF list ver. 1 - 6); 本村, 2021 (JAF list ver. 7 - 12); 本村, 2022 (JAF list ver. 13 - 18); 本村, 2023 (JAF list ver. 19 - 21)
Cobitis takatsuensis, Shikoku group
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
Ishihara et al., 2020Ishihara, M., Morita, K., Iguchi, Y., Takaku, K., Tanaka, K. & Kitagawa, T. 2020. Examination of unidentifiable spined loach individuals found in the overlap zone of two tetraploid species within a single river in Japan. Journal of Applied Ichthyology, 36(5): 668-674. DOI: 10.1111/jai.14081 URL
Nakajima et al., 2008Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. 2008. Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57: 10-15.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
biwae complex Tetraploid race
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
Tabata et al., 2016Tabata, R., Kakioka, R., Tominaga, K., Komiya, T. & Watanabe, K. 2016. Phylogeny and historical demography of endemic fishes in Lake Biwa: the ancient lake as a promoter of evolution and diversification of freshwater fishes in western Japan. Ecology and Evolution, DOI: 10.1002/ece3.2070 URL
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
biwae complex Diploid races
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
Miyazaki et al., 2014Miyazaki, Y., Nakae, M. & Senou, H. 2014. Ichthyofauna of the Kubo, Tochikura, and Ichinono river systems (Kitakami River drainage, northern Japan), with a comparison of predicted and surveyed species richness. Biodiversity Data Journal, 2: DOI: 10.3897/BDJ.2.e1093 URL
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
biwae complex Diploid races
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
biwae complex Diploid races
Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Cobitis striata (the Onga form of the middle race)
Nakajima et al., 2011Nakajima, J., Ohata, T. & Onikura, N. 2011. Longitudinal distribution patterns of three spined loach species (Cobitidae, Cobitis) in the Onga River system, northern Kyushu Island, Japan. Folia Zoologica, 60: 322–327.
Onga form of Cobitis stirata (middle race)
Nakajima et al., 2008Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. 2008. Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57: 10-15.
Onga form of middle race of Cobitis striata complex
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Nakajima et al., 2008Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. 2008. Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57: 10-15.
Hakata form of middle race of Cobitis striata complex
van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Tang et al., 2010Tang, K.L., Agnew, M.K., Hirt, M.V., Sado, T., Schneider, L.M., Freyhof, J., Sulaiman, Z., Swartz, E., Vidthayanon, C., Miya, M., Saitoh, K., Simons, A.M., Wood, R.M. & Mayden, R.L. 2010. Systematics of the subfamily Danioninae (Teleostei: Cypriniformes: Cyprinidae). Molecular Phylogenetics and Evolution, 57: 189-214.; Kottelat, 2012Kottelat, M. 2012. CONSPECTUS COBITIDUM: AN INVENTORY OF THE LOACHES OF THE WORLD (TELEOSTEI: CYPRINIFORMES: COBITOIDEI). THE RAFFLES BULLETIN OF ZOOLOGY, Supplement 26: 1–199.; Imoto et al., 2013Imoto, J.M., Saitoh, K., Sasaki, T., Yonezawa, T., Adachi, J., Kartavtsev, Y.P., Miya, M., Nishida, M. & Hanzawa, N. 2013. Phylogeny and biogeography of highly diverged freshwater fish species (Leuciscinae, Cyprinidae, Teleostei) inferred from mitochondrial genome analysis. Gene, 514: 112-124.; Perdices et al., 2016Perdices, A., Bohlen, J., Šlechtová, V. & Doadrio, I. 2016. Molecular Evidence for Multiple Origins of the European Spined Loaches (Teleostei, Cobitidae). PLoS ONE, 11(1): e0144628. DOI: 10.1371/journal.pone.0144628 URL; Schönhuth et al., 2018Schönhuth, S., Vukić, J., Šanda, R., Yang, L. & Mayden, R.L. 2018. Phylogenetic relationships and classification of the Holarctic family Leuciscidae (Cypriniformes: Cyprinoidei). Molecular Phylogenetics and Evolution, 127: 781-799. DOI: 10.1016/j.ympev.2018.06.026 URL
Nakajima et al., 2008Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. 2008. Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57: 10-15.
normal type (Setouchi form) of middle race in Cobitis striata complex
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL; Saitoh et al., 2010Saitoh, K., Chen, W.-J. & Mayden, R.L. 2010. Extensive hybridization and tetrapolyploidy in spined loach fish. Molecular Phylogenetics and Evolution, 56: 1001-1010.
Suzawa, 2006Suzawa, Y. 2006. A new loach, Cobitis shikokuensis (Teleostei: Cobitidae), from Shikoku Island, Japan. Ichthyological Research, 53: 315-322. DOI: 10.1007/s10228-006-0352-4 URL; Nakajima et al., 2008Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. 2008. Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57: 10-15.; Nakajima et al., 2011Nakajima, J., Ohata, T. & Onikura, N. 2011. Longitudinal distribution patterns of three spined loach species (Cobitidae, Cobitis) in the Onga River system, northern Kyushu Island, Japan. Folia Zoologica, 60: 322–327.; Kottelat, 2012Kottelat, M. 2012. CONSPECTUS COBITIDUM: AN INVENTORY OF THE LOACHES OF THE WORLD (TELEOSTEI: CYPRINIFORMES: COBITOIDEI). THE RAFFLES BULLETIN OF ZOOLOGY, Supplement 26: 1–199.; van der Laan, 2015van der Laan, R. 2015. Freshwater Fish List: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Eleventh edition. 910pp.; Perdices et al., 2016Perdices, A., Bohlen, J., Šlechtová, V. & Doadrio, I. 2016. Molecular Evidence for Multiple Origins of the European Spined Loaches (Teleostei, Cobitidae). PLoS ONE, 11(1): e0144628. DOI: 10.1371/journal.pone.0144628 URL; van der Laan, 2016van der Laan, R. 2016. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Seventeenth edition. 940pp.; Chen et al., 2017Chen, Y., He, D., Chen, H. & Chen, Y. 2017. Taxonomic study of the genus Niwaella (Cypriniformes: Cobitidae) from East China, with description of four new species. Zoological Systematics, 42(4): 490-507. DOI: 10.11865/zs.201723 URL; van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
Kitagawa et al., 2005Kitagawa, T., Jeon, S.-R., Kitagawa, E., Yoshioka, M., Kashiwagi, M. & Okazaki, T. 2005. Genetic relationships among the Japanese and Korean striated spined loach complex (Cobitidae: Cobitis) and their phylogenetic positions. Ichthyological Research, 52: 111-122. DOI: 10.1007/s10228-004-0261-3 URL
van der Laan, 2017van der Laan, R. 2017. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twentieth edition. 975pp.; van der Laan, 2018van der Laan, R. 2018. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-fourth edition. 1006pp.; van der Laan, 2019van der Laan, R. 2019. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-sixth edition. 1072pp.; Ishihara et al., 2020Ishihara, M., Morita, K., Iguchi, Y., Takaku, K., Tanaka, K. & Kitagawa, T. 2020. Examination of unidentifiable spined loach individuals found in the overlap zone of two tetraploid species within a single river in Japan. Journal of Applied Ichthyology, 36(5): 668-674. DOI: 10.1111/jai.14081 URL; van der Laan, 2020van der Laan, R. 2020. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Twenty-eighth edition. 1088pp.; van der Laan, 2021van der Laan, R. 2021. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-first Edition. 1102pp.; van der Laan, 2022van der Laan, R. 2022. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-fourth Edition. 1120pp.; van der Laan, 2023van der Laan, R. 2023. FRESHWATER FISH LIST: an alphabetic scientific name list of the world’s freshwater fishes and an overview of the scientific names used in the aquarium literature. Thirty-seventh Edition. 1130pp.
タンゴスジシマドジョウ
☆Nakajima, 2016Nakajima, J. 2016. Cobitis takenoi sp. n. (Cypriniformes, Cobitidae): a new spined loach from Honshu Island, Japan. Zookeys, 568: 119-128.; 中島・内山, 2017中島 淳・内山りゅう. 2017. 日本のドジョウ 形態・生態・文化と図鑑. 山と渓谷社,223pp. Amazon; 中坊編, 2018中坊徹次(編). 2018. 小学館の図鑑Z 日本魚類館. 小学館,xvi+524pp. Amazon; 細谷ほか, 2019細谷和海・内山りゅう・藤田朝彦・武内啓明・川瀬成吾. 2019. 山渓ハンディ図鑑15 増補改訂 日本の淡水魚. 山と渓谷社, 560pp. Amazon; 本村, 2020 (JAF list ver. 1 - 6); 本村, 2021 (JAF list ver. 7 - 12); 本村, 2022 (JAF list ver. 13 - 18); 本村, 2023 (JAF list ver. 19 - 21)
本種は異質4倍体の種で,mtDNAはチュウガタスジシマドジョウ,あるいはトサシマドジョウに近いことから(Ishihara et al. 2020),この2種の交雑に由来する可能性がある.形態的変異が大きいため,同所的に生息するオオシマドジョウと識別できないこともある.しかし,本種とオオシマドジョウの交雑は知られていない(Ishihara et al. 2020).
Ishihara, M., Morita, K., Iguchi, Y., Takaku, K., Tanaka, K. & Kitagawa, T. 2020. Examination of unidentifiable spined loach individuals found in the overlap zone of two tetraploid species within a single river in Japan. Journal of Applied Ichthyology, 36(5): 668-674. DOI: 10.1111/jai.14081 URL
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岡田彌一郎. 1938. 日本脊椎動物目録 A CATALOGUE OF VERTEBRATES OF JAPAN. 丸善株式会社, iv + 412pp. 国立国会図書館デジタルコレクション(参照2023-6-23). DOI: 10.11501/1680758 URL
Perdices, A., Bohlen, J., Šlechtová, V. & Doadrio, I. 2016. Molecular Evidence for Multiple Origins of the European Spined Loaches (Teleostei, Cobitidae). PLoS ONE, 11(1): e0144628. DOI: 10.1371/journal.pone.0144628 URL
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